You can set a specific node label here, which will be treated as a "magic" value during the evaluation of the restrictions on running jobs.

If a value is set here and you start a job that is an Inheritance Job and has restrictions on labels, Jenkins will check if you make any reference of the "magic" value in the label expression.

If not, Jenkins will add an "and" term with the negated magic value. That means, these jobs will not run on nodes that carry the magic label.
If there is a reference, Jenkins will do nothing and just return the label that you specified. This way, you can select manually whether or not to run jobs on test-machines that are marked with the "magic" label.